Fig. 1

Average values and associated variation for seven landscape metrics across six spatial extents. Grain was held constant [10 m] across all six spatial extents. The research was conducted in two pine savannas (Joseph W. Jones Ecological Research Center and Silver Lake Wildlife Management Area) located in southwestern Georgia, USA, 2011–2013. a Percent mature pine. b Percent mixed/pine hardwood. c Percent hardwood. d Percent young pine. e Percent agriculture. f Percent shrub/scrub. g Edge density