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Table 1 The relationship between more easily measured “soft functional traits” and less easily measured, but potentially more informative, “hard functional traits”

From: Functional perspectives on tropical tree demography and forest dynamics

Soft traits

Hard traits

Citations

Seed mass

• Seed persistence

• Germination requirements

• Survival to reproduction

• Dispersal mode

Díaz and Cabido (1997); Westoby et al. (2002); Cornelissen et al. (2003); Metz et al. (2010)

Wood density

• Aboveground storage of carbon

• Mechanical failure risk

• Diameter growth rate

• Mortality rate

• Timing of reproduction

• Hydraulic capacities

Carlquist (1975); Putz et al. (1983); Niklas (1997); Enquist et al. (1999); Cornelissen et al. (2003); van Gelder et al. (2006); Swenson and Enquist (2007)

Specific leaf area

• Mass-based maximum photosynthetic rate

• Potential relative growth rate

• Intrinsic water use efficiency

• Leaf palatability

• Leaf life span

• Leaf instantaneous photosynthetic rate

Blackman and Wilson (1951); Aerts (1990); Poorter and Remkes (1990); Lambers and Poorter (1992); Reich et al. (1992); Díaz and Cabido (1997); Reich et al. (1997); Reich et al. (1998); Diemer (1998); Lavorel and Garnier (2002); Westoby et al. (2002); Cornelissen et al. (2003); Elger and Willby (2003); Angert et al. (2007)

Leaf nitrogen content and leaf phosphorous content

• Mass-based maximum photosynthetic rate

• Nutritional quality

• Leaf instantaneous photosynthetic rate

Lavorel and Garnier (2002); Cornelissen et al. (2003)