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Table 1 Hierarchical organization of a typical vertebrate lifetime track (LiT)

From: A hierarchical path-segmentation movement ecology framework

Scale/segment

Time

Space

Categories

Data

Some approaches and methods

Constraint level: type of analysis

FuMEs\(^*\)

0.1–few secs

0–1000 cm

Stride, trot, run, twist, jump, flap

Accelerometer video

Machine learning\(^{*\dag }\), Newtonian mechanics

Highly constrained: biomechanical analyses

Meta- FuMEs\(^*\)

Resolution of relocation data\(^{*\ddag }\)

Average step length

Bivariate SL and TA distributions

SL and TA means, variances and correlations\(^{\dag \dag }\)

Cluster analysis ensemble filtering

Base level: movement ecology analyses

Short duration CAMs\(^\dag\)

0.1–10 min

A few meters to a few kms

Resting, browsing, riding thermals

secs to mins,

Path segmentation\(^{**}\)

Highly flexible: sub-hourly/hourly analysis

metaFuMEs and covariate data

Long duration CAMs\(^\dag\)

10–100 min

A few meters to many kms

Feeding traveling

mins to hours,

Highly flexible: sub-diel analysis

Short CAMs and Covariate data

DARs\(^\ddag\)

Fixed 24-h

Diel range

Central-place foraging Ranging

24 h of data CAM eequences

Periodograms (Fourier, wavelets)

Time constrained: diel cycle analysis

  

LiMPs\(^\S\)

Several days to a few months

Home ranges and their shifts over time

Philopatry, dispersal, migration, nomadic, recursion

DAR sequences

Clustering methods, ideal-free/despotic distribution analyses

Strongly constrained: lunar and seasonal cycle analyses

Partial or whole LiTs\(^\#\)

Paths encompassing several LiMPs

Habitat types

Territorial ranging, migratory nomadic

LiMP sequences

GIS toolbox, deep-learning\(^{*\dag }\), pattern anal.

Moderately constrained: question-at-hand analysis

  
  1. SL step-length, TA turning-angle
  2. The indicated spatiotemporal scaling applies best to most medium and large vertebrates, with faster/smaller scales needed for many smaller vertebrates, birds, and invertebrates.)
  3. \(^*\)Fundamental movement elements; \(^\dag\)canonical activity modes; \(^\ddag\)diel activity routines
  4. \(^\S\)Lifetime movement phases; \(^\#\)lifetime tracks
  5. \(^{*\dag }\)Machine/deep learning methods can be applied at any scale but may be particularly useful here
  6. \(^{*\ddag }\)Around 5 s to 1 min
  7. \(^{**}\)Includes hidden Markov models (HMMs) and behavioral change-point analyses (BCPA)
  8. \(^{\dag \dag }\)See “Stochastic walk statistics” section in main text